Normative models of enhancer function
R. Grah, B. Zoller, G. Tkadik, BioRxiv (2020).

Nonequilibrium models of optimal enhancer function
R. Grah, B. Zoller, G. Tkadik, Proceedings of the National Academy of Sciences of the United States of America 117 (2020) 31614–31622.

Gene regulation across scales – how biophysical constraints shape evolution
R. Grah, Gene Regulation across Scales – How Biophysical Constraints Shape Evolution, Institute of Science and Technology Austria, 2020.

Gene amplification as a form of population-level gene expression regulation
I. Tomanek, R. Grah, M. Lagator, A.M.C. Andersson, J.P. Bollback, G. Tkadik, C.C. Guet, Nature Ecology &; Evolution 4 (2020) 612–625.